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- The seed can be in the centre of the wing, like the elms and the hoptree.
- The seed can be on one side, with the wing extending to the other side, making the seed self rotate it falls.
- botanical fruit = seed with covering tissue, but not a fruit in ordinary speech.
- Fergus, Charles (2002). Trees of Pennsylvania and the Northeast. Stackpole Books. . https://books.google.com/books?id=4LKBcxNzHZAC.
- Catalogues of Plants: no. 1. Arthur J.C; Coulter J.M. and Barnes C.R. 1881. Catalogue of the phaenogamous and vascular cryptogamous plants of Indiana.. https://books.google.com/books?id=ds5HAQAAMAAJ.
- Transactions of the Royal Scottish Arboricultural Society. Douglas & Foulis. 1881. https://books.google.com/books?id=Z57kAAAAMAAJ.
- Alexander, David E.; Vogel, Steven (2004). Nature's Flyers: Birds, Insects, and the Biomechanics of Flight. JHU Press. . https://books.google.com/books?id=zj395mz_GYkC.
- Ralph Lorenz 2006. Spinning flight: dynamics of frisbees, boomerangs, samaras and skipping stones, , Copernicus New York. ISBN 0-387-30779-6
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