Prokaryote

Structure of a prokaryotic bacteria cell

Prokaryotes (or Monera) are some of the simplest living things.^[1] They are unicellular organisms and they include two major divisions of simple living beings: bacteria, and Archaea.^[2] They generally do not have a cell nucleus, nuclear membrane or cell organelles, however a small number of exceptions have been found. Prokaryotes possess genetic material in the form of a single DNA loop called a Nucleoid that floats around freely inside the cell. Bacteria include the Cyanobacteria (formerly known as blue-green algae due to their photosynthetic capacity).

Some form biofilms that are somewhat like multicellular organisms.

Besides a nucleus, prokaryotes lack other things eukaryotes (cells with a true nucleus) have. They reproduce without fusion of gametes. They do not have membranes. This means that there are no vacuoles, Golgi apparatus, endoplasmic reticula or other organelles inside the cell. However, they do have ribosomes, though of a simpler kind than eukaryotes. Eukaryote cells include organelles which were once free-living prokaryotes.

Taxonomy

In 1977, Carl Woese proposed dividing prokaryotes into the Bacteria and Archaea (originally Eubacteria and Archaebacteria) because of the major differences in the structure and genetics between the two groups of organisms. This arrangement of Eukaryota (also called "Eukarya"), Bacteria, and Archaea is called the three-domain system, replacing the traditional two-empire system.^[3]

The Archaea include simple organisms which were first discovered in extreme environments. Most of them can survive at very high or very low temperatures. Some of them can also survive in highly salty, acidic or alkaline water. Some have been found in geysers, black smokers or oil wells.

The prokaryotes may not be a monophyletic, taxonomic group of common descent, but merely a group of similar organisms. The relationships between Archaea, Eubacteria and Eukaryota are assessed differently by different scientists. The three-domain system of Carl Woese treats them as one monophyletic group.

Prokaryote Media

Diagram of a typical prokaryotic cell
3D animation of a prokaryotic cell that shows all the elements that it is composed of
Phylogenetic ring showing the diversity of prokaryotes, and symbiogenetic origins of eukaryotes
Phylogenetic and symbiogenetic tree of living organisms, showing the origins of eukaryotes and prokaryotes
A Novel Representation Of The Tree Of Life
Diagram of the origin of life with the Eukaryotes appearing early, not derived from Prokaryotes, as proposed by Richard Egel in 2012. This view, one of many on the relative positions of Prokaryotes and Eukaryotes, implies that the universal common ancestor was relatively large and complex.
Comparison of eukaryotes vs. prokaryotes
Phylogenetic tree showing the diversity of prokaryotes.*This 2018 proposal shows eukaryotes emerging from the archaean Asgard group which represents a modern version of the eocyte hypothesis. Unlike earlier assumptions, the division between bacteria and the rest is the most important difference between organisms.