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The Santana Formation is a high quality fossil site in Brazil. It is in northeastern Brazil's Arar ipe Basin. It qualifies as a Lagerstätte, which means it is one of the best fossil sites, with many undisturbed fossils.
Some 25 species of fish are found, many with stomach contents preserved. This lets palaeontologists study predator–prey relationships in the ecosystem. There are also fine examples of pterosaurs, reptiles, amphibians, invertebrates (especially insects) and plants. Even dinosaurs are found here: a new maniraptor was described in 1996.
Limestone nodules formed around dead organisms; this preserved even the soft parts of their anatomy. The limestone was etched away using a mild acid. The fossils were then preserved by the 'transfer technique' which means they were embedded in a plastic resin such as polyester or epoxy.
The nearby Crato Formation is also a lagerstätte. It was produced in similar conditions, about 10 million years before the Santana Formation.
Various remains of non-avian theropods, avialans, ornithischians, and possibly oviraptorosaurs have been found in Estado do Ceara, Brazil. Examples are spinosaurs like Irritator and maniraptors like Santanaraptor.
The pterosaurs are all pterodactyls, because the long-tailed rhamphorhynchoidea died out at the Jurassic–Cretaceous junction. Ornithocheirus and Tupuxuara are examples. The inland sea also had turtles, such as Santanachelys.
- Maisey J.G. et al. 1991. Laboratory preparation techniques. In Maisey J.G. (ed) Santana fossils: an illustrated atlas. Tfh Pubns, 99–103. ISBN 0-86622-549-8
- Weishampel, David B et al. 2004. Dinosaur distribution (early Cretaceous, South America). In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds): The Dinosauria. 2nd ed, Berkeley: University of California Press, 563-570. ISBN 0-520-24209-2